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[1] 13/APR/26

Dynamical Aspects of Pattern Formation

May 18 -- May 20, 2026


Venue:    [May 18 & 19] Aoba Science Hall,    [May 20] Kawai Hall,
                Graduate School of Science, Tohoku University
                Aobayama, Aoba-ku, Sendai    
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May 18         May 19         May 20


Abstracts


Monday, May 18, 2026

9:20     Opening


9:30 – 10:20   Anna Marciniak-Czochra (Heidelberg University)
Selecting patterns through mechanics: Insights from a new PDE model

Abstract: In this talk, I will present a mechanochemical, PDE-based framework motivated by recent experiments on regenerating epithelia, with Hydra as a model system. The framework couples reaction-diffusion dynamics of a morphogen with elasticity equations governing tissue deformation. A positive feedback between mechanical strain and morphogen production drives symmetry breaking and leads to the emergence of stable, single-peaked steady states. Using bifurcation and stability analysis, I will characterise the onset of pattern formation and identify parameter regimes in which these solutions exist and remain robust. This mechanism provides a mathematically distinct route to effective long-range inhibition, in contrast to curvature-morphogen coupling or classical Turing-type diffusion-driven instabilities. Model predictions are in quantitative agreement with experimental observations, suggesting that mechanically mediated feedback may constitute a fundamental organising principle in epithelial pattern formation.

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10:30 – 11:20   Kousuke Kuto (Waseda University)
Bifurcation and classification in cross-diffusion limits of the SKT model

Abstract: This talk concerns asymptotic regimes of positive steady states in the Shigesada-Kawasaki-Teramoto competition model under large cross-diffusion limits. We present a systematic classification of limiting regimes according to unilateral or full cross-diffusion limits, together with boundary conditions including Neumann and Dirichlet cases. We explain how these four settings lead to distinct limiting shadow systems and determine qualitative structures of coexistence states, such as segregation, extinction, or concentration profiles. This classification provides a unified framework that organizes previously known results and clarifies structural relations between different limiting procedures. This talk is based on joint work with Profs. Yaping Wu (Capital Normal University) and Toru Kan (Osaka Metropolitan University).
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11:30 -- 12:00   Szymon Cygan (Heidelberg University & University of Wroclaw)
Bifurcation analysis and oscillations in mechanochemical model of pattern formation

Abstract: In this talk, I will present a nonlocal reaction-diffusion model coupling morphogen dynamics with tissue mechanics to describe pattern formation in regenerating Hydra spheroids. The model is given by the integro-differential equation $\partial_t u = D \partial_{xx} u - u + \kappa \frac{e^u}{\int_0^1 e^u\, dx}$, for which I will perform a rigorous bifurcation analysis of the homogeneous steady state and show that symmetry breaking occurs via pitchfork bifurcations, which are supercritical for $\kappa > 1.5$ and subcritical for $\kappa < 1.5$. In the subcritical regime, I will further identify fold bifurcations leading to bistability between homogeneous and heterogeneous states.
I will then extend the framework to a time-dependent domain by introducing a tissue length $\ell(t)$, and show that the qualitative features of these oscillations are determined by the underlying bifurcation structures, in particular by whether the system undergoes subcritical or supercritical bifurcations.   (
pdf file)
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lunch break

13:30 -- 14:20   Hirokazu Ninomiya (Meiji University)
Dynamics of Curves under the Anisotropic Area-Preserving Curvature Flows

Abstract: In 1986, Gage studied area-preserving curvature flows in a two-dimensional homogeneous medium and proved that an initially convex closed curve remains convex and converges to a circle as time approaches infinity. However, in many applications, such as cell motility and the motion of active matter, the medium is often inhomogeneous. In this talk, I will introduce the following area-preserving curvature flow in an inhomogeneous medium, which can be regarded as a natural extension of the curvature flow in a homogeneous medium. Through this extension, a closed curve numerically moves towards the higher medium by the flow. To show this fact, the case is considered in which the area enclosed by the closed curve is small. In this situation, it is shown that the dynamics of its center is approximated by a gradient flow determined by the distribution of the inhomogeneous medium and that its center moves toward the critical point of the inhomogeneous medium.

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14:30 -- 15:20   Kentaro Fujie (Tohoku University)
Entropy production structure of Keller–Segel systems

Abstact: In seminal works by Keller and Segel, some mathematical model was introduced to describe aggregation phenomena and pattern formations induced by chemotaxis effect. In the mathematical analysis of large time behaviour of solutions to this model, its entropy structure plays an important role. In this talk, we focus on the entropy production (the time derivative of the entropy) of the Keller–Segel system and related parabolic equations. The time evolution of the entropy production is discussed and some application of this property is also given.

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15:40 -- 16:30   Goro Akagi
Optimal rate of convergence to non-isolated asymptotic profiles for fast-diffusion in thin annuli

Abstract: In this talk, we discuss the optimal rate of convergence to asymptotic profiles of solutions, which vanish in finite time, to the Fast Diffusion Equation posed on bounded domains. Quantitative studies in this direction have developed significantly since the celebrated work of Bonforte and Figalli (2021). However, in most existing results, asymptotic profiles are assumed to be nondegenerate, meaning that the linearized operator has a trivial kernel. In fact, such nondegeneracy holds for generic domains. On the other hand, it is well known that least-energy solutions to the Emden--Fowler equation in thin annuli are nonradial. As a consequence, they are degenerate and thus fall outside the scope of previous works. In this talk, inspired by recent work of König and Yu (2024$^+$), we establish the optimal convergence rate to such degenerate asymptotic profiles for certain space dimensions. This talk is based on a recent joint work with Norihisa Ikoma (Keio University) and Yasunori Maekawa (Kyoto University).   (pdf file)


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16:40 -- 17:10   Rong Lei (Tohoku University)
Homogenization of Stokes–Cahn–Hilliard system with logarithmic free energy in porous media

Abstract: We investigate the homogenization of a phase-field model for two-phase flow in porous media, incorporating surface tension effects. The model is governed by the coupled Stokes–Cahn–Hilliard system with a logarithmic free energy, where the two fluids are separated by a diffuse interface of finite width. We first prove some uniform energy estimate. Then, by the method of two-scale convergence and unfolding theory, we derive the homogenized limit, which takes the form of a generalized Richards’ equation. This is joint work with Jun Masamune.


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17:30 -- 19:00     Welcome



Monday, May 19, 2026


9:30 -- 10:20   Chiun-Chuan Chen (National Taiwan University)
Propagation direction in the two species Lotka-Volterra competition system with diffusion

Abstract: We investigate the propagation direction of bistable traveling waves in the two-species Lotka–Volterra competition–diffusion model under strong competition. From an ecological perspective, the sign of the wave speed is critical, as it dictates which species eventually prevails. We propose a mini-max characterization of the coefficients in the reaction terms when the wave speed is zero. Applying our characterization, some explicit conditions under which the speed sign can be determined are obtained. We also focus on a near-symmetric scenario where intrinsic growth rates and inter-specific competition coefficients are identical, leaving diffusion rates as the sole source of asymmetry. This framework is motivated by Girardin et al.’s conjecture, which proposes that the species with the higher diffusion rate gains a competitive advantage, directly dictating the wave speed's sign. In this talk, we explore the subtle regime where inter-specific competition weakens, approaching the strong-competition boundary.

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10:30 -- 11:20   Yasuhito Miyamoto (University of Tokyo)
Exact solutions describing very slow layer oscillations in a shadow reaction-diffusion system

Abstract: We show in a rigorous way that a stable internal single-layer stationary solution is destabilized by the Hopf bifurcation as the time constant exceeds a certain critical value. Moreover, the exact critical value and the exact period of oscillatory solutions can be obtained. The exact period indicates that the oscillation is very slow, i.e., the period is of order O(e^{C/ε}) We also rigorously prove that Hopf bifurcations from multi-layer stationary solutions occur. In this case anti-phase horizontal oscillations of layers are shown by formal calculations. Numerical experiments show that the exact period agrees with the numerical period of a nearly periodic solution near the Hopf bifurcation point. Anti-phase (out of phase) horizontal oscillations of layers are numerically observed.

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11:30 -- 12:00   Shun-Chieh Wang
The Non-Monotone Waves in the Two-Species Lotka-Volterra Competition Model and Application

Abstract: We investigate the Lotka–Volterra competitive reaction-diffusion equation with a focus on the co-existence phenomenon. To understand the system's dynamics, we investigate traveling wave solutions. Our goal is to find solutions connecting the two equilibria $(0,0)$ and $(u^*,v^*)$. We consider the following system \begin{equation} \begin{cases} u''-su'+u(1-u-cv)=0, \ \xi \in \mathbb{R},\\ dv''-sv'+v(a-bu-v)=0,\\ (u,v)(-\infty)=(0,0), \ (u,v)(+\infty)=(u^*,v^*), \end{cases} \end{equation} under the weak competition condition; b < a < \frac{1}{c}.
Previous studies have established the existence of strictly monotone solution; however, the solution is not unique.
In this work, we successfully construct the non-monotone solution. We also use this solution to construct the front-pulse solution.   (
pdf file)

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lunch break

13:30 -- 14:20   Lorenzo Cavallina (Tohoku University)
On the geometry of solutions to the two-phase Serrin problem

Abstract: In this talk, I will discuss an overdetermined problem arising in the shape optimization problem of maximizing the torsional rigidity of composite beams made of two distinct materials separated by an interface. I will focus on two-phase configurations for which this overdetermined problem admits a solution, and I will present several qualitative and geometric properties of such configurations. In particular, I will explain how the geometry of the interface affects the overall configuration's shape, and show when optimal configurations are radially symmetric and when they are not.

14:30 -- 15:20   Shinya Okabe (Tohoku University)
Length penalised ideal curve flow for planar closed curves

Abstract: The ideal functional is defined by the Dirichlet energy of the curvature of curves and originates from an elasticity problem posed by Jacob Bernoulli in 1694. In this talk, we consider the length-penalised ideal curve flow for planar closed curves. Although one might expect the functional to have a similar variational structure to that of the well-known elastic energy due to its origin, the ideal functional has a quite different structure. One of the main results of this talk is that the length-penalised ideal curve flow blows up in finite time under a suitable condition on the initial curves. Furthermore, there are infinitely many kinds of critical points of the ideal functional with length penalisation. We will state this precisely as a second main result of the talk. This talk is based on joint work with Glen Wheeler at the University of Wollongong.
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15:40 -- 16:30   Kanako Suzuki (Ibaraki University)
Existence of a traveling wave solution induced by the basic production term in an activator–inhibitor system

Abstract: We study the effect of a basal production term on an activator–inhibitor type reaction–diffusion system. When the basal production is a positive constant, the system admits a unique positive constant steady state. Under an appropriate change of variables, another constant steady state induced by the basal production term emerges. We discuss the existence of traveling wave solutions connecting this new steady state and the positive constant steady state. This is a joint work with Izumi Takagi (Tohoku University).
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16:30 -- 18:30    Poster Session

TBA ( University)   
TBA

Abstract: TBA

TBA ( University)   
TBA

Abstract: TBA



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Wednesday, May 20, 2026



9:30 -- 10:00   Finn Münnich (Heidelberg University)
Emergent patterns in systems coupling diffusive and non-diffusive components

Abstract: Reaction-diffusion-ODE systems have emerged as powerful models for biological pattern formation, capturing the interplay between diffusive and non-diffusive nonlinear processes. These systems exhibit a rich variety of spatial structures, including classical Turing patterns and far-from-equilibrium patterns exhibiting jumps. Previous studies have primarily focused on diffusion-driven instability (DDI) generated by instability of the purely non-diffusive subsystem, which destabilizes classical Turing patterns. In contrast, we show that DDI can also arise from subsystems that couple non-diffusive and slowly diffusing components, leading to dynamics involving three distinct spatial scales. Moreover, we prove the existence of far-from-equilibrium patterns in a general reaction-diffusion-ODE framework. As illustrative examples, we apply our results to specific models, demonstrating the broad applicability of the approach.

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10:10 -- 11:00   Izumi Takagi (Tohoku University)
Looking back on half a century of the research on pattern formation in reaction-diffusion systems.

Abstract: It has been more than fifty years since the activator-inhibitor model was proposed by Gierer and Meinhardt, which stimulated mathematical studies on qualitative properties of solutions to nonlinear partial differential equations. In this talk I would like to sketch the (very personalized) history of studies on reaction-diffusion equations and related fields based on what I witnessed.

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11:15 -- 12:30   Group Discussions


12:35   Closing


Organizing Committee: Goro Akagi (Sendai), Anna Marciniak-Czochra (Heidelberg), Kanako Suzuki (Mito), Izumi Takagi (Sendai)

Supported in part by
    JSPS Grant-in-Aid for Scientific Research (A) #24H00184 "Evolution equations describing singularities of non-equilibrium systems beyond the linear-response regime and development of nonlocal-nonlinear analysis" (Akagi)
    JSPS Grant-in-Aid for Scientific Research (C) #23K03177 "On the role of forcing terms in the dynamics of reaction-diffusion-ODE systems" (Suzuki)
    JSPS Grant-in-Aid for Scientific Research (C) #23K03176 "Behaviour of Nonstationary Solutions to Reaction-Diffusion Systems Possessing Continua of Stationary Solutions” (Takagi)
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created on 6/apr/26